Carcharodus flocciferus (Zeller, 1847)


Carcharodus flocciferus: Female (S-Germany, Bad Kohlgrub, 27. June 2021) Carcharodus flocciferus: Female (S-Germany, Bad Kohlgrub, 27. June 2021) Carcharodus flocciferus: Female (S-Germany, Bad Kohlgrub, 27. June 2021) Carcharodus flocciferus: Female (S-Germany, Bad Kohlgrub, 27. June 2021) Carcharodus flocciferus: Female (S-Germany, Bad Kohlgrub, 27. June 2021) Carcharodus flocciferus: Female (S-Germany, Bad Kohlgrub, 27. June 2021) Carcharodus flocciferus: Female, sleeping position (S-Germany, Bad Kohlgrub, 27. June 2021) Carcharodus flocciferus: Female, sleeping position (S-Germany, Bad Kohlgrub, 27. June 2021) Carcharodus flocciferus: Female (S-Germany, Bad Kohlgrub, 13. July 2020) Carcharodus flocciferus: Male (SE-France, Col d Carcharodus flocciferus: Female Carcharodus flocciferus: Female Carcharodus flocciferus: Adult upper side [S] Carcharodus flocciferus: Adult (Col de Larche, France, 2100m above sea level, July 2012) [N] Carcharodus flocciferus: Adult lower side [S] Carcharodus flocciferus: Mating behaviour (S-Germany, Bad Kohlgrub, 13. July 2020) [N] Carcharodus flocciferus: Egg (S-Germany, Bad Kohlgrub, 13. July 2020) [N] Carcharodus flocciferus: Egg (S-Germany, Bad Kohlgrub, 13. July 2020) [N] Carcharodus flocciferus: Egg (S-Germany, Bad Kohlgrub, 13. July 2020) [N] Carcharodus flocciferus: Egg (S-Germany, Bad Kohlgrub, 13. July 2020) [N] Carcharodus flocciferus: Egg (S-Germany, Bad Kohlgrub, 13. July 2020) [N] Carcharodus flocciferus: Egg on Betonica pradica (SE-France, Col d Carcharodus flocciferus: Egg on Betonica pradica (SE-France, Col d Carcharodus flocciferus: Ovum [N] Carcharodus flocciferus: Ovum [N] Carcharodus flocciferus: Larva in the first instar (SE-France, Col d Carcharodus flocciferus: Larva in the first instar (SE-France, Col d Carcharodus flocciferus: Larva in the first instar (S-Germany, Bad Kohlgrub, 13. July 2020) Carcharodus flocciferus: Larva in the first instar (S-Germany, Bad Kohlgrub, 13. July 2020) Carcharodus flocciferus: Larva in the first instar (SE-France, Col d Carcharodus flocciferus: Larva in the second instar (S-Germany, Bad Kohlgrub, August 2020) Carcharodus flocciferus: Larva in the second instar (SE-France, Col d Carcharodus flocciferus: Larva in the third instar (SE-France, Col d Carcharodus flocciferus: Larva in the third instar prior to hibernation (SE-France, Col d Carcharodus flocciferus: Hibernation shelter of a larva in the third instar (SE-France, Col d Carcharodus flocciferus: Half-grown larva (S-Germany, Bad Kohlgrub, August 2020) Carcharodus flocciferus: Half-grown larva (S-Germany, Bad Kohlgrub, August 2020) Carcharodus flocciferus: Half-grown larva (S-Germany, Bad Kohlgrub, August 2020) Carcharodus flocciferus: Young larva (late August 2012, S-Bavaria) [M] Carcharodus flocciferus: Half-grown larva [M] Carcharodus flocciferus: Half-grown larva [M] Carcharodus flocciferus: Half-grown larva prior to hibernation (S-Germany, Bad Kohlgrub, late August 2020) Carcharodus flocciferus: Half-grown larva after hibernation (S-Germany, Bad Kohlgrub, 10. April 2021) Carcharodus flocciferus: Larva in the spring Carcharodus flocciferus: Larva in the penultimate instar (S-Germany, Bad Kohlgrub, May 2021) Carcharodus flocciferus: Larva [M] Carcharodus flocciferus: Larva [S] Carcharodus flocciferus: Larva [S] Carcharodus flocciferus: Larva [S] Carcharodus flocciferus: Larva Carcharodus flocciferus: Larva Carcharodus flocciferus: Larva [N] Carcharodus flocciferus: Larva in the final instar (S-Germany, Bad Kohlgrub, May 2021) Carcharodus flocciferus: Larva in the final instar (S-Germany, Bad Kohlgrub, May 2021) Carcharodus flocciferus: Larva in the final instar (S-Germany, Bad Kohlgrub, May 2021) Carcharodus flocciferus: Larva in the final instar (S-Germany, Bad Kohlgrub, May 2021) Carcharodus flocciferus: Larva in the final instar (S-Germany, Bad Kohlgrub, May 2021) Carcharodus flocciferus: Larva in the final instar (S-Germany, Bad Kohlgrub, May 2021) Carcharodus flocciferus: Larva in the final instar (S-Germany, Bad Kohlgrub, May 2021) Carcharodus flocciferus: Larva in the final instar (S-Germany, Bad Kohlgrub, May 2021) Carcharodus flocciferus: Larva in the final instar (S-Germany, Bad Kohlgrub, May 2021) Carcharodus flocciferus: Larva in the final instar (S-Germany, Bad Kohlgrub, May 2021) Carcharodus flocciferus: Pupa [M] Carcharodus flocciferus: Pupa Carcharodus flocciferus: Pupa (S-Germany, Bad Kohlgrub, June 2021) Carcharodus flocciferus: Pupa (S-Germany, Bad Kohlgrub, June 2021) Carcharodus flocciferus: Pupa (S-Germany, Bad Kohlgrub, June 2021) Carcharodus flocciferus: Pupa (S-Germany, Bad Kohlgrub, June 2021) Carcharodus flocciferus: Pupa (S-Germany, Bad Kohlgrub, June 2021) Carcharodus flocciferus: Oviposition habitat with Betonica pradica (SE-France, Col d Carcharodus flocciferus: Hostplant of the SW-Alps: Betonica pradica (SE-France, Col d Carcharodus flocciferus: Hostplant of the SW-Alps: Betonica pradica (SE-France, Col d Carcharodus flocciferus: Habitat (SE-France, Col d Carcharodus flocciferus: Oviposition habitat (S-Germany, Bad Kohlgrub, 13. July 2020) [N] Carcharodus flocciferus: Oviposition habitat (S-Germany, Bad Kohlgrub, 13. July 2020) [N] Carcharodus flocciferus: Oviposition habitat (S-Germany, Bad Kohlgrub, 13. July 2020) Carcharodus flocciferus: Betonica officinalis with larval shelter (S-Bavaria, late August 2012) [N] Carcharodus flocciferus: Young larval shelter (S-Bavaria, late August 2012) [N] Carcharodus flocciferus: Shelter of a larva prior to hibernation (S-Germany, Bad Kohlgrub, August 2021) Carcharodus flocciferus: Shelter of a larva prior to hibernation (S-Germany, Bad Kohlgrub, August 2021) Carcharodus flocciferus: Shelter of a larva prior to hibernation (S-Germany, Bad Kohlgrub, August 2021) Carcharodus flocciferus: Shelter of a half-grown larva after hibernation (S-Germany, Bad Kohlgrub, 10. April 2021) Carcharodus flocciferus: Larval shelter (S-Germany, Bad Kohlgrub, May 2021) Carcharodus flocciferus: Larval shelter (S-Germany, Bad Kohlgrub, May 2021) Carcharodus flocciferus: Larval shelter (S-Germany, Bad Kohlgrub, May 2021) Carcharodus flocciferus: Pupal shelter (S-Germany, Bad Kohlgrub, May 2021) Carcharodus flocciferus: Habitat in the northern pre-Alps [N] Carcharodus flocciferus: Larval habitat in a wetland in S-Bavaria. The meadow in the foreground has been mowed too early and too low on the ground. Thus I still observed only a few surviving larvae in the small embankment in the left half of the image. Late August 2012. [N] Carcharodus flocciferus: Habitat (S-Germany, Bad Kohlgrub, 13. July 2020) [N]

Host plants:
The larva most often develops on Stachis (= Betonica) officinalis. In the higher altitudes of the southwestern Alps the caterpillars live on Stachys (Betonica) pradica (= hirsuta), which is very similar to Stachys officinalis.

Habitat:
In Central Europe Carcharodus flocciferus lives in wetland complexes (fens) with nutrient-poor, low growing and drier areas. The insect still occurs in Germany only at the foothills of the Alps from Lake Constance to Lake Chiemsee.
In the Southwestern Alps Carcharodus flocciferus is apparently more common at higher altitudes (well correlated with the distribution of Betonica pradica). I found the butterflies at 1600 - 2200m above sea level both in the Italian Alps near Cuneo and at almost every pass in the French Alpes-De-Haute-Provence and the Hautes-Alpes. At the Col de Var numerous ovipositions on Betonica pradica on low growing, drier and raised spots were observed within mucky areas as it is often also the case in Central European fens. In the northern Alps Carcharodus flocciferus does not occur in the true mountain region above 1200m above sea level. Here also lacks Betonica officinalis. Betonica pradica occurs more in the Southern and especially Southwestern Alps.

Life cycle:
In general a single generation flies in June and July. Occasionally a very partial second generation appears in August/September. Eggs are laid on the upper leaf surface of small, mostly sterile plants of Betonica officinalis, where the caterpillars create small housings. Here they partially gnaw the leaf stem, so that the leaf bags are often characterized by wilting and are easily noticeable. Hibernation occurs as half-grown larva. In the spring the mature caterpillars can be found mainly in May. Sometimes concentrations of larvae (each singly in its own shelter) occur within the wetlands at elevated, low growing places that are more reminiscent of dry grassland sites. Pupation takes place in a webbed shelter between leaves at the food plant.

Endangerment factors:
This species is threatened with extinction in Central Europe. Previously the main causes were mainly reforestation, reducing of site size (intersection with corn fields) and intensification of wet meadows. Today it is more the eutrophication of the remainders, felting, bush encroachment, road construction, improper maintenance measures, radical ditch clearances, increasing isolation etc. Even today sometimes roads are built through the last precious fens.

In the southwestern Alps Carcharodus flocciferus is still more widespread. Threats are there the intense tourism and its infrastructure measures and too intense grazement on the one hand, but also abandonment and felting of the vegetation on the other.

Remarks:
Carcharodus flocciferus is distributed from Morocco (very local) across central and southern Europe (here especially in mountainous areas) and spreads to the Altai.



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