Coenonympha leander (Esper, [1784])


Coenonympha leander: Female (Vitsi, Florina, Greece, 1500m asl, late June 2013) [N] Coenonympha leander: Female (Vitsi, Florina, Greece, 1500m asl, late June 2013) [N] Coenonympha leander: Female (Vitsi, Florina, Greece, 1500m asl, late June 2013) [N] Coenonympha leander: Female (Vitsi, Florina, Greece, 1500m asl, late June 2013) [N] Coenonympha leander: Female (Vitsi, Florina, Greece, 1500m asl, late June 2013) [N] Coenonympha leander: Female (Vitsi, Florina, Greece, 1500m asl, late June 2013) [N] Coenonympha leander: Female (Vitsi, Florina, Greece, 1500m asl, late June 2013) [N] Coenonympha leander: Female (Vitsi, Florina, Greece, 1500m asl, late June 2013) [N] Coenonympha leander: Male (e.o. Vitsi 2013/2014) [S] Coenonympha leander: Male (Vitsi, Florina, Greece, 1500m asl, late June 2013) [N] Coenonympha leander: Male (Vitsi, Florina, Greece, 1500m asl, late June 2013) [N] Coenonympha leander: Female at oviposition (Vitsi, Florina, Greece, 1500m asl, late June 2013) [N] Coenonympha leander: Fresh egg (Vitsi, late June 2013) [M] Coenonympha leander: L1-larva (e.o. Vitsi) [S] Coenonympha leander: L1-larva (e.o. Vitsi) [S] Coenonympha leander: L1-larva at the end of this instar (e.o. Vitsi) [S] Coenonympha leander: Young larva (e.o. Vitsi 2013) [S] Coenonympha leander: Young larva (e.o. Vitsi 2013) [S] Coenonympha leander: Larva in the hibernation instar (e.o. Vitsi 2013) [S] Coenonympha leander: Larva in the hibernation instar (e.o. Vitsi 2013) [S] Coenonympha leander: Larva in penultimate instar after hibernation (e.o. N-Greece, Vitsi N Kastoria, 2013/2014) [S] Coenonympha leander: Larva in penultimate instar after hibernation (e.o. N-Greece, Vitsi N Kastoria, 2013/2014) [S] Coenonympha leander: Larva in last instar (e.o. N-Greece, Vitsi N Kastoria, 2013/2014) [S] Coenonympha leander: Larva in last instar (e.o. N-Greece, Vitsi N Kastoria, 2013/2014) [S] Coenonympha leander: Larva in last instar (e.o. N-Greece, Vitsi N Kastoria, 2013/2014) [S] Coenonympha leander: Larva in last instar (e.o. N-Greece, Vitsi N Kastoria, 2013/2014) [S] Coenonympha leander: Larva in last instar (e.o. N-Greece, Vitsi N Kastoria, 2013/2014) [S] Coenonympha leander: Larva in last instar (e.o. N-Greece, Vitsi N Kastoria, 2013/2014) [S] Coenonympha leander: Larva in last instar (e.o. N-Greece, Vitsi N Kastoria, 2013/2014) [S] Coenonympha leander: Larva in last instar (e.o. N-Greece, Vitsi N Kastoria, 2013/2014) [S] Coenonympha leander: Larva in last instar (e.o. N-Greece, Vitsi N Kastoria, 2013/2014) [S] Coenonympha leander: Larva in last instar (e.o. N-Greece, Vitsi N Kastoria, 2013/2014) [S] Coenonympha leander: Larva in last instar (e.o. N-Greece, Vitsi N Kastoria, 2013/2014) Coenonympha leander: Larva in last instar (e.o. N-Greece, Vitsi N Kastoria, 2013/2014) [S] Coenonympha leander: Larva in last instar (e.o. N-Greece, Vitsi N Kastoria, 2013/2014) [S] Coenonympha leander: Larva in last instar (e.o. N-Greece, Vitsi N Kastoria, 2013/2014) [S] Coenonympha leander: Larva in last instar (e.o. N-Greece, Vitsi N Kastoria, 2013/2014) [S] Coenonympha leander: Larva in last instar (e.o. N-Greece, Vitsi N Kastoria, 2013/2014) [S] Coenonympha leander: Pupa (e.o. N-Greece, Vitsi N Kastoria, 2013/2014) [S] Coenonympha leander: Pupa (e.o. N-Greece, Vitsi N Kastoria, 2013/2014) [S] Coenonympha leander: Oviposition habitat (Vitsi, 1500m asl, late June 2013) [N] Coenonympha leander: Habitat on mount Vitsi in 1500m asl (Greece, late June 2013) [N] Coenonympha leander: Habitat on mount Vitsi in 1500m asl (Greece, late June 2013) [N] Coenonympha leander: Habitat on mount Vitsi in 1500m asl (Greece, late June 2013) [N]

Host plants:
The larvae feed on grasses. I observed oviposition on a blue-green, fine-leaved tussock grass of the Festuca ovina group in Northern Greece.

Habitat:
Coenonympha leander inhabits nutrient-poor grasslands, grassy slopes and pastures especially near the woodland or near to scrub.

Life cycle:
The adults occur between mid-May and July (according to altitude). I observed worn males and rather fresh females on mount Vitsi (Florina) in Northern Greece in about 1500m asl in late June and early July 2013. For oviposition the females landed on the Festuca tussocks, crawled further down and attached the egg near ground-level to grass leaves.
The adults often sit on the ground or on leaves of bushes and small trees up to several meters in height in order to sunbath with closed wings. I watched most individuals directly on the edge of beech woodland to grasslands.

According to breedeing results the larva hibernates half-grown in ground-level parts of the tussocks and should be mature in late April or May.

Endangerment factors:
Coenonympha leander is probably endangered in lower altitudes because of ever increasing intensity of agriculture.

Remarks:
Coenonympha leander occurs in Europe in parts of the Balkan peninsula from Northern Greece to Romania and Moldavia. It is further recorded in southern Russia and parts of Asia.



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