Cucullia celsiae (Herrich-Schäffer, [1850])


Cucullia celsiae: Adult (Cyprus, Paphos forest, Ano Panagia, at light, legit Franz Grünwald, late February 2018) [S] Cucullia celsiae: Adult (Cyprus, Paphos forest, Ano Panagia, at light, legit Franz Grünwald, late February 2018) [S] Cucullia celsiae: Adult (Cyprus, Paphos forest, Ano Panagia, at light, legit Franz Grünwald, late February 2018) [S] Cucullia celsiae: Adult (Cyprus, Paphos forest, Ano Panagia, at light, legit Franz Grünwald, late February 2018) [S] Cucullia celsiae: Adult (Cyprus, Paphos forest, Ano Panagia, at light, legit Franz Grünwald, late February 2018) [S] Cucullia celsiae: Adult (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018, moth hatched in late January 2019) [S] Cucullia celsiae: Adult (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018, moth hatched in late January 2019) [S] Cucullia celsiae: Adult (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018, moth hatched in late January 2019) [S] Cucullia celsiae: Adult (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018, moth hatched in late January 2019) [S] Cucullia celsiae: Adult (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018, moth hatched in late January 2019) [S] Cucullia celsiae: Adult (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018, moth hatched in late January 2019) [S] Cucullia celsiae: Adult (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018, moth hatched in late January 2019) [S] Cucullia celsiae: Adult (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018, moth hatched in late January 2019) [S] Cucullia celsiae: Adult (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018, moth hatched in late January 2019) [S] Cucullia celsiae: Adult (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018, moth hatched in late January 2019) [S] Cucullia celsiae: Adult (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018, moth hatched in late January 2019) [S] Cucullia celsiae: Egg (W-Cyprus, Paphos forest, early April 2018) [M] Cucullia celsiae: Egg (W-Cyprus, Paphos forest, early April 2018) [M] Cucullia celsiae: Eggs (W-Cyprus, Paphos forest, early April 2018) [M] Cucullia celsiae: Eggs (e.l. rearing, W-Cyprus, Paphos forest, larva in February 2018) [M] Cucullia celsiae: Egg (W-Cyprus, Paphos forest, early April 2018) [M] Cucullia celsiae: Egg some days after oviposition (W-Cyprus, Paphos forest, early April 2018) [M] Cucullia celsiae: Egg, becoming light grey prior to hatching (W-Cyprus, Paphos forest, early April 2018) [M] Cucullia celsiae: Egg, becoming light grey prior to hatching (W-Cyprus, Paphos forest, early April 2018) [M] Cucullia celsiae: Egg shell after hatching (W-Cyprus, Paphos forest, early April 2018) [M] Cucullia celsiae: L1-larva (e.o.-rearing, W-Cyprus, Paphos forest, eggs found in early April 2018) [S] Cucullia celsiae: L1-larva (W-Cyprus, Paphos forest, early April 2018) [M] Cucullia celsiae: L2-larva (W-Cyprus, Paphos forest, early April 2018) [S] Cucullia celsiae: L2-larva (W-Cyprus, Paphos forest, early April 2018) [S] Cucullia celsiae: L2-larva (W-Cyprus, Paphos forest, early April 2018) [M] Cucullia celsiae: L2-larva (W-Cyprus, Paphos forest, early April 2018) [M] Cucullia celsiae: L2-larva (W-Cyprus, Paphos forest, early April 2018) [M] Cucullia celsiae: L3-larva (W-Cyprus, Paphos forest, early April 2018) [M] Cucullia celsiae: L3-larva (W-Cyprus, Paphos forest, early April 2018) [M] Cucullia celsiae: L3-larva (W-Cyprus, Paphos forest, early April 2018) [M] Cucullia celsiae: L3-larva (W-Cyprus, Paphos forest, early April 2018) [M] Cucullia celsiae: L3-larva (W-Cyprus, Paphos forest, early April 2018) [M] Cucullia celsiae: L3-larva (W-Cyprus, Paphos forest, early April 2018) [M] Cucullia celsiae: L4-larva (W-Cyprus, Paphos forest, early April 2018) [N] Cucullia celsiae: L4-larva (W-Cyprus, Paphos forest, early April 2018) [N] Cucullia celsiae: L4-larva (W-Cyprus, Paphos forest, early April 2018) [M] Cucullia celsiae: L4-larva (W-Cyprus, Paphos forest, early April 2018) [M] Cucullia celsiae: L4-larva (W-Cyprus, Paphos forest, early April 2018) [M] Cucullia celsiae: L4-larva (W-Cyprus, Paphos forest, early April 2018) [M] Cucullia celsiae: L4-larva (W-Cyprus, Paphos forest, early April 2018) [M] Cucullia celsiae: Larva in the last instar (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018) [S] Cucullia celsiae: Larva in the last instar (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018) [S] Cucullia celsiae: Larva in the last instar (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018) [S] Cucullia celsiae: Larva in the last instar (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018) [S] Cucullia celsiae: Larva in the last instar (W-Cyprus, Agios Therapon, early April 2018) [M] Cucullia celsiae: Larva in the last instar (W-Cyprus, Agios Therapon, early April 2018) [M] Cucullia celsiae: Larva in the last instar (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018) [S] Cucullia celsiae: Larva in the last instar (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018) [S] Cucullia celsiae: Larva in the last instar (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018) [S] Cucullia celsiae: Larva in the last instar (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018) [S] Cucullia celsiae: Larva in the last instar (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018) [S] Cucullia celsiae: Larva in the last instar (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018) [S] Cucullia celsiae: Larva in the last instar (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018) [S] Cucullia celsiae: Larva in the last instar (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018) [S] Cucullia celsiae: Larva in the last instar (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018) [S] Cucullia celsiae: Larva in the last instar (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018) [S] Cucullia celsiae: Larva in the last instar (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018) [S] Cucullia celsiae: Larva in the last instar (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018) [S] Cucullia celsiae: Larva in the last instar (W-Cyprus, Agios Therapon, early April 2018) [M] Cucullia celsiae: Larva in the last instar (W-Cyprus, Agios Therapon, early April 2018) [M] Cucullia celsiae: Larva in the last instar (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018) [S] Cucullia celsiae: Larva in the last instar (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018) [S] Cucullia celsiae: Larva in the last instar (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018) [S] Cucullia celsiae: Larva in the last instar (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018) [S] Cucullia celsiae: Larva in the last instar (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018) [S] Cucullia celsiae: Larva in the last instar (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018) [S] Cucullia celsiae: Larva in the last instar (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018) [S] Cucullia celsiae: Larva in the last instar (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018) [S] Cucullia celsiae: Larva in the last instar (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018) [S] Cucullia celsiae: Larva in the last instar (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018) [S] Cucullia celsiae: Larva in the last instar (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018) [S] Cucullia celsiae: Larva in the last instar (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018) [S] Cucullia celsiae: Larva in the last instar (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018) [S] Cucullia celsiae: Larva in the last instar (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018) [S] Cucullia celsiae: Larva in the last instar (W-Cyprus, Agios Therapon, early April 2018) [M] Cucullia celsiae: Larva in the last instar (W-Cyprus, Agios Therapon, early April 2018) [M] Cucullia celsiae: Larva in the last instar (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018) [S] Cucullia celsiae: Larva in the last instar (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018) [S] Cucullia celsiae: Larva in the last instar (W-Cyprus, Agios Therapon, early April 2018) [M] Cucullia celsiae: Larva in the last instar (W-Cyprus, Agios Therapon, early April 2018) [M] Cucullia celsiae: Larva searchin for a pupation site (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018) [S] Cucullia celsiae: Parasitized, dead larva with pupated parasitoid inside (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018) [S] Cucullia celsiae: Cocoon (e.l. rearing, W-Cyprus, Agios Therapon, larva in early April 2018) [S] Cucullia celsiae: Pupa (e.l. rearing, W-Cyprus, Agios Therapon, larva in early April 2018) [S] Cucullia celsiae: Pupa (e.l. rearing, W-Cyprus, Agios Therapon, larva in early April 2018) [S] Cucullia celsiae: Pupa (e.l. rearing, W-Cyprus, Agios Therapon, larva in early April 2018) [S] Cucullia celsiae: Pupa (e.l. rearing, W-Cyprus, Agios Therapon, larva in early April 2018) [S] Cucullia celsiae: Pupa (e.l. rearing, W-Cyprus, Agios Therapon, larva in early April 2018) [S] Cucullia celsiae: Pupa, already partially developed prior to hibernation (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018) [S] Cucullia celsiae: Pupa, already partially developed prior to hibernation (e.l. rearing, W-Cyprus, Paphos forest, larva in early April 2018) [S] Cucullia celsiae: Larval habitat with Scutellaria cypria (W-Cyprus, Paphos forest, early April 2018) [N] Cucullia celsiae: Larval habitat with Scutellaria cypria (W-Cyprus, Paphos forest, early April 2018) [N] Cucullia celsiae: Larval habitat with Scutellaria cypria (W-Cyprus, Paphos forest, early April 2018) [N] Cucullia celsiae: Larval habitat with Scutellaria cypria (W-Cyprus, Paphos forest, early April 2018) [N] Cucullia celsiae: Larval habitat with Scutellaria cypria (W-Cyprus, Paphos forest, early April 2018) [N] Cucullia celsiae: Larval habitat with Scutellaria cypria (W-Cyprus, Paphos forest, early April 2018) [N] Cucullia celsiae: Larval habitat with Scutellaria cypria (W-Cyprus, Paphos forest, early April 2018) [N] Cucullia celsiae: Larval habitat with Scutellaria cypria (W-Cyprus, Paphos forest, early April 2018) [N] Cucullia celsiae: Larval habitat with Scutellaria cypria (W-Cyprus, Paphos forest, early April 2018) [N] Cucullia celsiae: Larval habitat with Scutellaria cypria (W-Cyprus, Paphos forest, early April 2018) [N] Cucullia celsiae: Larval habitat with Scutellaria cypria (W-Cyprus, Paphos forest, early April 2018) [N] Cucullia celsiae: Larval habitat with Scutellaria cypria (W-Cyprus, Paphos forest, early April 2018) [N] Cucullia celsiae: Larval habitat with Scutellaria cypria (W-Cyprus, Paphos forest, early April 2018) [N] Cucullia celsiae: Larval habitat with Scutellaria cypria (W-Cyprus, Paphos forest, early April 2018) [N] Cucullia celsiae: Larval habitat with Scutellaria cypria (W-Cyprus, Paphos forest, early April 2018) [N] Cucullia celsiae: Larval habitat with Scutellaria cypria (W-Cyprus, Paphos forest, early April 2018) [N] Cucullia celsiae: Larval habitat with Scutellaria cypria (W-Cyprus, Paphos forest, early April 2018) [N] Cucullia celsiae: Larval habitat with Scutellaria cypria (W-Cyprus, Paphos forest, early April 2018) [N] Cucullia celsiae: Larval habitat with Scutellaria cypria (W-Cyprus, Paphos forest, early April 2018) [N] Cucullia celsiae: Larval habitat with Scutellaria cypria (W-Cyprus, Paphos forest, early April 2018) [N] Cucullia celsiae: Larval habitat with Scutellaria cypria (W-Cyprus, Paphos forest, early April 2018) [N] Cucullia celsiae: Larval habitat in a limestone region with Scutellaria cypria (W-Cyprus, Agios Therapon, early April 2018) [N] Cucullia celsiae: Larval habitat in a limestone region with Scutellaria cypria (W-Cyprus, Agios Therapon, early April 2018) [N] Cucullia celsiae: Larval habitat in a limestone region with Scutellaria cypria (W-Cyprus, Agios Therapon, early April 2018) [N] Cucullia celsiae: Larval habitat in a limestone region with Scutellaria cypria (W-Cyprus, Agios Therapon, early April 2018) [N] Cucullia celsiae: Larval habitat in a limestone region with Scutellaria cypria (W-Cyprus, Agios Therapon, early April 2018) [N] Cucullia celsiae: Larval habitat in a limestone region with Scutellaria cypria (W-Cyprus, Agios Therapon, early April 2018) [N] Cucullia celsiae: Larval habitat in a limestone region with Scutellaria cypria (W-Cyprus, Agios Therapon, early April 2018) [N]

Host plants:
In Cyprus, the larvae feed on Scutellaria cypria (Lamiaceae). Thus it is expected to live on related Scutelaria species in other regions, too. These larvae did not accept Brassicaceae species in captivity. Thus the reputed record on Hesperum desertorum (Brassicaceae) in literature is certainly wrong.

Habitat:
Cucullia celsiae obviously prefers rocky places or at least sites with open soil and screes with the host plants. The host plants also occur in garigues, open shrubland, grasslands, woodland edges, open pine woodland and mountain slopes which expands the habitat spectrum. Spots of open soil seem to be important.

Life cycle:
The pupa hibernates partly developed and the moths occur between February (e.g. Cyprus February to April), but most often not until March to May, in higher altitudes even later. The larvae develop between late March/April to July, in Greek lowlands usually between mid-April and early June.

Remarks:
Cucullia celsiae occurs from the Southern Balkans (e.g. Albania, Croatia, Romania, Greece, Macedonia, Bulgaria) across Asia Minor, Cyprus and the Levant to Iran.

Literature:
Wagner, W. (2018): The mystery of the preimaginal stages of Cucullia celsiae Herrich-Schäffer, [1850] solved in Cyprus, and second record of Cucullia gozmanyi (G. & L. Ronkay, 1994) in Greece(Lepidoptera: Noctuidae).- Nachr. entomol. Ver. Apollo, N. F. 39 (2): 69-74.



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